Genealogies in multitype populations: branching processes and structured coalescents
Abstract.
This talk focuses on the interplay between type and ancestry in two different multitype population models. In the first part, we briefly discuss the long-term behavior of critical multitype branching processes conditioned on survival, both with respect to the forward and the ancestral processes. Despite substantial differences in forward-time behavior and required techniques, their ancestral processes retain key structural similarities to the supercritical case. The main part of the talk then focuses on structured populations divided into $d$ colonies, where individuals migrate at rates proportional to a global scaling parameter $K$. We sample $N(K)$ individuals evenly across colonies and trace their ancestral lineages backward in time. Within each colony, coalescence occurs at a constant rate as in the Kingman coalescent. We encode the system's state as a $d$-dimensional vector of empirical measures, recording both current lineage locations and the colonies of their sampled descendants. Our focus is on how the sample size affects the asymptotic behavior of this process as $K \to \infty$ (representing fast migration), distinguishing two regimes: the critical-sampling regime ($N(K) \sim K$) and the large-sampling regime ($N(K) \gg K$). After suitable time-space rescaling, we prove convergence to $d$-dimensional coagulation equations in both sampling regimes. In the critical regime, the solution admits a representation via a multitype birth-death process; in the large-sample regime, via the entrance law of a multitype Feller diffusion.